About the Authors

Jianying Li

Contributed equally to this work with: Jianying Li, Qihui Wang

Affiliation Zhejiang Key Laboratory for Organogenesis and Regenerative technology, Institute of Developmental and Regenerative Biology, College of Life and Environmental Sciences, Hangzhou Normal University, Hangzhou 310036, China

Qihui Wang

Contributed equally to this work with: Jianying Li, Qihui Wang

Affiliation Zhejiang Key Laboratory for Organogenesis and Regenerative technology, Institute of Developmental and Regenerative Biology, College of Life and Environmental Sciences, Hangzhou Normal University, Hangzhou 310036, China

Ying Cui

Affiliation Zhejiang Key Laboratory for Organogenesis and Regenerative technology, Institute of Developmental and Regenerative Biology, College of Life and Environmental Sciences, Hangzhou Normal University, Hangzhou 310036, China

Xueqin Yang

Affiliation Zhejiang Key Laboratory for Organogenesis and Regenerative technology, Institute of Developmental and Regenerative Biology, College of Life and Environmental Sciences, Hangzhou Normal University, Hangzhou 310036, China

Yan Li

Affiliation Zhejiang Key Laboratory for Organogenesis and Regenerative technology, Institute of Developmental and Regenerative Biology, College of Life and Environmental Sciences, Hangzhou Normal University, Hangzhou 310036, China

Xiaoyun Zhang

Affiliation Zhejiang Key Laboratory for Organogenesis and Regenerative technology, Institute of Developmental and Regenerative Biology, College of Life and Environmental Sciences, Hangzhou Normal University, Hangzhou 310036, China

Mengsheng Qiu

Affiliations Zhejiang Key Laboratory for Organogenesis and Regenerative technology, Institute of Developmental and Regenerative Biology, College of Life and Environmental Sciences, Hangzhou Normal University, Hangzhou 310036, China, Department of Anatomical Sciences and Neurobiology, University of Louisville, Louisville, Kentucky 40292, United States of America

Ze Zhang

Affiliation Department of Ophthalmology, Tulane University Medical Center, New Orleans, Louisiana 70112, United States of America

Zunyi Zhang

zunyi.zhang@idrbio.org

Affiliation Zhejiang Key Laboratory for Organogenesis and Regenerative technology, Institute of Developmental and Regenerative Biology, College of Life and Environmental Sciences, Hangzhou Normal University, Hangzhou 310036, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ZYZ JYL QHW ZZ XYZ. Performed the experiments: JYL QHW YC XQY YL XYZ. Analyzed the data: JYL QHW XQY YL XYZ ZZ ZYZ. Contributed reagents/materials/analysis tools: JYL MSQ ZZ QHW ZYZ. Wrote the paper: ZYZ JYL ZZ.