About the Authors

Andrew M. Gormley

Affiliation Landcare Research, Lincoln, New Zealand

David M. Forsyth

dave.forsyth@delwp.vic.gov.au

Current Address: Arthur Rylah Institute for Environmental Research, Department of Environment, Land, Water and Planning, Heidelberg, Victoria, Australia

Affiliation Science and Capability Group, Department of Conservation, Christchurch, New Zealand

Elaine F. Wright

Affiliation Science and Capability Group, Department of Conservation, Christchurch, New Zealand

John Lyall

Affiliation Department of Conservation, Hokitika, New Zealand

Mike Elliott

Affiliation Department of Conservation, Hokitika, New Zealand

Mark Martini

Affiliation Department of Conservation, Hokitika, New Zealand

Benno Kappers

Affiliation Science and Capability Group, Department of Conservation, Wellington, New Zealand

Mike Perry

Affiliation Landcare Research, Palmerston North, New Zealand

Meredith McKay

Affiliation Science and Capability Group, Department of Conservation, Christchurch, New Zealand

Competing Interests

The authors have following competing interest: AMG and MP were employees of Landcare Research, a commercial company. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: AMG DMF EFW JL MP M. Mckay. Performed the experiments: EFW JL ME M. Mckay MP M. Martini. Analyzed the data: AMG DMF. Wrote the paper: AMG DMF. Coordinated data management: BK.