About the Authors

Masaya Watanabe

Contributed equally to this work with: Masaya Watanabe, Takashi Ueda

Affiliation Department of Anatomy and Neuroscience, Graduate School of Medical Sciences, Nagoya City University, Nagoya, Japan

Takashi Ueda

Contributed equally to this work with: Masaya Watanabe, Takashi Ueda

tueda@med.nagoya-cu.ac.jp

Affiliation Department of Anatomy and Neuroscience, Graduate School of Medical Sciences, Nagoya City University, Nagoya, Japan

Yasuhiro Shibata

Affiliation Department of Anatomy and Neuroscience, Graduate School of Medical Sciences, Nagoya City University, Nagoya, Japan

Natsuko Kumamoto

Affiliation Department of Anatomy and Neuroscience, Graduate School of Medical Sciences, Nagoya City University, Nagoya, Japan

Shoichi Shimada

Affiliation Department of Neuroscience and Cell Biology, Osaka University Graduate School of Medicine, Suita, Japan

Shinya Ugawa

Affiliation Department of Anatomy and Neuroscience, Graduate School of Medical Sciences, Nagoya City University, Nagoya, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MW TU. Performed the experiments: MW TU YS NK. Analyzed the data: MW TU YS SS. Contributed reagents/materials/analysis tools: MW TU. Wrote the paper: MW TU SS SU.