About the Authors

Yoshiko Nishiyama

Affiliation Department of Psychiatry and Neurosciences, Graduate School of Biomedical and Health Sciences, Hiroshima University, Hiroshima, Japan

Yasumasa Okamoto

Affiliation Department of Psychiatry and Neurosciences, Graduate School of Biomedical and Health Sciences, Hiroshima University, Hiroshima, Japan

Yoshihiko Kunisato

Affiliation Department of Psychology, School of Human Sciences, Senshu University, Kawasaki, Japan

Go Okada

Affiliation Department of Psychiatry and Neurosciences, Graduate School of Biomedical and Health Sciences, Hiroshima University, Hiroshima, Japan

Shinpei Yoshimura

Affiliation Department of Psychology, Faculty of Psychology, Otemon Gakuin University, Osaka, Japan

Yoshihiro Kanai

Affiliation Department of Human Science, Faculty of Liberal Arts, Tohoku Gakuin University, Sendai, Japan

Takanao Yamamura

Affiliation Department of Psychiatry and Neurosciences, Graduate School of Biomedical and Health Sciences, Hiroshima University, Hiroshima, Japan

Atsuo Yoshino

Affiliation Department of Psychiatry and Neurosciences, Graduate School of Biomedical and Health Sciences, Hiroshima University, Hiroshima, Japan

Ran Jinnin

Affiliation Department of Psychiatry and Neurosciences, Graduate School of Biomedical and Health Sciences, Hiroshima University, Hiroshima, Japan

Koki Takagaki

Affiliation Department of Psychiatry and Neurosciences, Graduate School of Biomedical and Health Sciences, Hiroshima University, Hiroshima, Japan

Keiichi Onoda

Affiliation Department of Neurology, Shimane University, Izumo, Japan

Shigeto Yamawaki

yamawaki@hiroshima-u.ac.jp

Affiliation Department of Psychiatry and Neurosciences, Graduate School of Biomedical and Health Sciences, Hiroshima University, Hiroshima, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YN YO Y. Kunisato GO S. Yoshimura KO S. Yamawaki. Performed the experiments: YN Y. Kunisato S. Yoshimura Y. Kanai TY AY RJ. Analyzed the data: YN Y. Kunisato GO S. Yoshimura KT KO. Wrote the paper: YN YO Y. Kunisato GO S. Yoshimura Y. Kanai KO S. Yamawaki.