About the Authors

Yusuke Fukuda

yusuke.fukuda@nt.gov.au

Affiliation Northern Territory Department of Land Resource Management, Palmerston, Northern Territory, Australia

Charlie Manolis

Contributed equally to this work with: Charlie Manolis, Keith Saalfeld, Alain Zuur

Affiliation Wildlife Management International Pty. Limited, Karama, Northern Territory, Australia

Keith Saalfeld

Contributed equally to this work with: Charlie Manolis, Keith Saalfeld, Alain Zuur

Affiliation Northern Territory Department of Land Resource Management, Palmerston, Northern Territory, Australia

Alain Zuur

Contributed equally to this work with: Charlie Manolis, Keith Saalfeld, Alain Zuur

Affiliation Highland Statistics Limited, Newburgh, United Kingdom

Competing Interests

CM is an employee of Wildlife Management International Pty., Limited. AZ is an employee of Highland Statistics Limited. There are no patents, products in development, or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: YF AZ. Performed the experiments: YF. Analyzed the data: YF CM KS AZ. Contributed reagents/materials/analysis tools: YF CM AZ. Wrote the paper: YF CM KS AZ.