About the Authors

Ju Rong

Contributed equally to this work with: Ju Rong, Qiuxia Li

Affiliation Division of Rheumatology, the Third Affiliated Hospital of Sun Yat-sen University, Tianhe Road 600, Guangzhou 510630, China

Qiuxia Li

Contributed equally to this work with: Ju Rong, Qiuxia Li

Affiliation Division of Rheumatology, the Third Affiliated Hospital of Sun Yat-sen University, Tianhe Road 600, Guangzhou 510630, China

Pingping Zhang

Affiliation Division of Rheumatology, the Third Affiliated Hospital of Sun Yat-sen University, Tianhe Road 600, Guangzhou 510630, China

Xinyu Wu

Affiliation Division of Rheumatology, the Third Affiliated Hospital of Sun Yat-sen University, Tianhe Road 600, Guangzhou 510630, China

Jinxian Huang

Affiliation Division of Rheumatology, the Third Affiliated Hospital of Sun Yat-sen University, Tianhe Road 600, Guangzhou 510630, China

Chao Li

Affiliation Division of Rheumatology, the Third Affiliated Hospital of Sun Yat-sen University, Tianhe Road 600, Guangzhou 510630, China

Zetao Liao

Affiliation Division of Rheumatology, the Third Affiliated Hospital of Sun Yat-sen University, Tianhe Road 600, Guangzhou 510630, China

Yingying Xie

Affiliation Division of Rheumatology, the Third Affiliated Hospital of Sun Yat-sen University, Tianhe Road 600, Guangzhou 510630, China

Qing Lv

Affiliation Division of Rheumatology, the Third Affiliated Hospital of Sun Yat-sen University, Tianhe Road 600, Guangzhou 510630, China

Qiujing Wei

Affiliation Division of Rheumatology, the Third Affiliated Hospital of Sun Yat-sen University, Tianhe Road 600, Guangzhou 510630, China

Tianwang Li

Affiliation Division of Rheumatology, the Third Affiliated Hospital of Sun Yat-sen University, Tianhe Road 600, Guangzhou 510630, China

Jianlin Huang

Affiliation Division of Rheumatology, the Third Affiliated Hospital of Sun Yat-sen University, Tianhe Road 600, Guangzhou 510630, China

Shuangyan Cao

Affiliation Division of Rheumatology, the Third Affiliated Hospital of Sun Yat-sen University, Tianhe Road 600, Guangzhou 510630, China

Yan Shen

Affiliation The Institute of Basic Medical Sciences, Chinese Academy of Medical Sciences & Peking Union Medical College and Chinese National Human Genome Research Center, Beijing, China

Jieruo Gu

gujieruo@163.com

Affiliation Division of Rheumatology, the Third Affiliated Hospital of Sun Yat-sen University, Tianhe Road 600, Guangzhou 510630, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JRG. Performed the experiments: JR QXL PPZ XYW JXH CL ZTL YYX QL. Analyzed the data: JR QXL PPZ XYW JXH CL ZTL YYX. Contributed reagents/materials/analysis tools: QJW TWL JLH SYC. Wrote the paper: JR PPZ XYW JXH ZTL. Revised it critically for important intellectual content: YS.