About the Authors
- Hirotoshi Sato
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* E-mail: h-sato@sys.bot.kyoto-u.ac.jp
Affiliation Center for Ecological Research, Kyoto University 509–3, 2-chome, Hirano, Otsu, Shiga, 520–2113 Japan
- Akifumi S. Tanabe
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Affiliations National Research Institute of Fisheries Science, Fisheries Research Agency, 2-12-4 Fukuura, Kanazawa-ku, Yokohama, Kanagawa, 236–8648 Japan, Graduate School of Global Environmental Studies, Kyoto University, Yoshida-nihonmatsu-cho, Sakyo-ku, Kyoto, Kyoto, 606–8501, Japan
- Hirokazu Toju
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Affiliation Graduate School of Human and Environmental Studies, Kyoto University, Yoshida-nihonmatsu-cho, Sakyoku Kyoto, 606–8501 Japan
Competing Interests
The authors have declared that no competing interests exist.
Author Contributions
Conceived and designed the experiments: HS. Performed the experiments: HS HT. Analyzed the data: HS AT. Contributed reagents/materials/analysis tools: HS AT HT. Wrote the paper: HS.