About the Authors

Christina R. Savage

Affiliation Department of Microbiology, Immunology, and Molecular Genetics, University of Kentucky College of Medicine, Lexington, Kentucky, United States of America

William K. Arnold

Affiliation Department of Microbiology, Immunology, and Molecular Genetics, University of Kentucky College of Medicine, Lexington, Kentucky, United States of America

Alexandra Gjevre-Nail

Affiliation Department of Microbiology, Immunology, and Molecular Genetics, University of Kentucky College of Medicine, Lexington, Kentucky, United States of America

Benjamin J. Koestler

Affiliation Department of Microbiology and Molecular Genetics, Michigan State University, East Lansing, Michigan, United States of America

Eric L. Bruger

Affiliation Department of Microbiology and Molecular Genetics, Michigan State University, East Lansing, Michigan, United States of America

Jeffrey R. Barker

Affiliation Department of Molecular Genetics and Microbiology, Center for Microbial Pathogenesis, Duke University, Durham, North Carolina, United States of America

Christopher M. Waters

Affiliation Department of Microbiology and Molecular Genetics, Michigan State University, East Lansing, Michigan, United States of America

Brian Stevenson

brian.stevenson@uky.edu

Affiliation Department of Microbiology, Immunology, and Molecular Genetics, University of Kentucky College of Medicine, Lexington, Kentucky, United States of America

Competing Interests

BS is an Academic Editor of PLoS One. The authors declare that no other competing interests exist.

Author Contributions

Conceived and designed the experiments: CRS JRB CMW BS. Performed the experiments: CRS WKA AG-N BJK ELB. Analyzed the data: CRS WKA BJK ELB CMW BS. Contributed reagents/materials/analysis tools: JRB CMW BS. Wrote the paper: CRS BS.