About the Authors

Shanshan Liu

Affiliations Key Lab of Animal Epidemiology and Zoonosis of Ministry of Agriculture, College of Veterinary Medicine, China Agricultural University,Beijing 100193, China, Institute of Animal Husbandry and Veterinary Medicine, Beijing Academy of Agricultural and Forestry Sciences, Beijing 100097, China

Wei Sun

Affiliation Institute of Animal Husbandry and Veterinary Medicine, Beijing Academy of Agricultural and Forestry Sciences, Beijing 100097, China

Jun Chu

Affiliation Key Lab of Animal Epidemiology and Zoonosis of Ministry of Agriculture, College of Veterinary Medicine, China Agricultural University,Beijing 100193, China

Xiufen Huang

Affiliation Institute of Animal Husbandry and Veterinary Medicine, Beijing Academy of Agricultural and Forestry Sciences, Beijing 100097, China

Zongxue Wu

Affiliation Key Lab of Animal Epidemiology and Zoonosis of Ministry of Agriculture, College of Veterinary Medicine, China Agricultural University,Beijing 100193, China

Minxin Yan

Affiliation Institute of Animal Husbandry and Veterinary Medicine, Beijing Academy of Agricultural and Forestry Sciences, Beijing 100097, China

Qiang Zhang

Affiliation Key Lab of Animal Epidemiology and Zoonosis of Ministry of Agriculture, College of Veterinary Medicine, China Agricultural University,Beijing 100193, China

Peng Zhao

Affiliation Key Lab of Animal Epidemiology and Zoonosis of Ministry of Agriculture, College of Veterinary Medicine, China Agricultural University,Beijing 100193, China

Joseph U. Igietseme

Affiliation National Center for Emerging and Zoonotic Infectious Diseases, CDC, Atlanta, Georgia, 30333, United States of America

Carolyn M. Black

Affiliation National Center for Emerging and Zoonotic Infectious Diseases, CDC, Atlanta, Georgia, 30333, United States of America

Cheng He

hecheng@cau.edu.cn (CH); liyngqng@yahoo.com (YQL)

Affiliation Key Lab of Animal Epidemiology and Zoonosis of Ministry of Agriculture, College of Veterinary Medicine, China Agricultural University,Beijing 100193, China

Yongqing Li

hecheng@cau.edu.cn (CH); liyngqng@yahoo.com (YQL)

Affiliation Institute of Animal Husbandry and Veterinary Medicine, Beijing Academy of Agricultural and Forestry Sciences, Beijing 100097, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SL CH YL. Performed the experiments: SL WS XH JC ZW MY QZ PZ. Analyzed the data: SL WS. Contributed reagents/materials/analysis tools: YL CH XH. Wrote the paper: SL WS JUI CMB CH YL.