About the Authors

Sarah F. Harpenslager

S.Harpenslager@science.ru.nl

Affiliation Department of Aquatic Ecology and Environmental Biology, Institute for Water and Wetland Research, Radboud University, Nijmegen, The Netherlands

Alfons J. P. Smolders

Affiliations Department of Aquatic Ecology and Environmental Biology, Institute for Water and Wetland Research, Radboud University, Nijmegen, The Netherlands, B-Ware Research Centre, Nijmegen, The Netherlands

Ariët A. M. Kieskamp

Affiliation Department of Aquatic Ecology and Environmental Biology, Institute for Water and Wetland Research, Radboud University, Nijmegen, The Netherlands

Jan G. M. Roelofs

Affiliation Department of Aquatic Ecology and Environmental Biology, Institute for Water and Wetland Research, Radboud University, Nijmegen, The Netherlands

Leon P. M. Lamers

Affiliation Department of Aquatic Ecology and Environmental Biology, Institute for Water and Wetland Research, Radboud University, Nijmegen, The Netherlands

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SH AS AK JR LL. Performed the experiments: SH AK. Analyzed the data: SH AK. Wrote the paper: SH AS AK JR LL.