About the Authors

Akiko Iizuka-Kogo

akogo@gunma-u.ac.jp

Affiliations Department of Anatomy I, Fujita Health University School of Medicine, Aichi, Japan, Department of Anatomy and Cell Biology, Gunma University Graduate School of Medicine, Gunma, Japan

Takao Senda

Affiliations Department of Anatomy I, Fujita Health University School of Medicine, Aichi, Japan, Department of Anatomy, Gifu University Graduate School of Medicine, Gifu, Japan

Tetsu Akiyama

Affiliation Laboratory of Molecular and Genetic Information, Institute of Molecular and Cellular Biosciences, The University of Tokyo, Tokyo, Japan

Atsushi Shimomura

Affiliations Department of Anatomy I, Fujita Health University School of Medicine, Aichi, Japan, The Department of Communication Disorders, School of Psychological Science, Health Sciences University of Hokkaido, Hokkaido, Japan

Ryuji Nomura

Affiliation Department of Anatomy I, Fujita Health University School of Medicine, Aichi, Japan

Yoshimi Hasegawa

Affiliation Department of Anatomy I, Fujita Health University School of Medicine, Aichi, Japan

Ken-ichi Yamamura

Affiliation Division of Developmental Genetics, Institute of Resource Development Analysis, Kumamoto University, Kumamoto, Japan

Hiroshi Kogo

Affiliation Department of Anatomy and Cell Biology, Gunma University Graduate School of Medicine, Gunma, Japan

Nobuhiko Sawai

Affiliation Department of Anatomy and Cell Biology, Gunma University Graduate School of Medicine, Gunma, Japan

Toshiyuki Matsuzaki

Affiliation Department of Anatomy and Cell Biology, Gunma University Graduate School of Medicine, Gunma, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AIK TS. Performed the experiments: AIK. Analyzed the data: AIK AS RN. Contributed reagents/materials/analysis tools: TA YH KY NS TM. Wrote the paper: AIK HK.