About the Authors

Zhen-Zhen Wu

Affiliations State Key Laboratory of Tropical Oceanography and Key Laboratory of Marine Bio-resources Sustainable Utilization, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou, China, Haikou Marine Environmental Monitoring Central Station, State Oceanic Administration, Haikou, China

Zhi-Wei Che

Affiliation Haikou Marine Environmental Monitoring Central Station, State Oceanic Administration, Haikou, China

You-Shao Wang

Affiliation State Key Laboratory of Tropical Oceanography and Key Laboratory of Marine Bio-resources Sustainable Utilization, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou, China

Jun-De Dong

Affiliations State Key Laboratory of Tropical Oceanography and Key Laboratory of Marine Bio-resources Sustainable Utilization, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou, China, Tropical Marine Biological Research Station in Hainan, Chinese Academy of Sciences, Sanya, China

Mei-Lin Wu

mlwu@scsio.ac.cn

Affiliation State Key Laboratory of Tropical Oceanography and Key Laboratory of Marine Bio-resources Sustainable Utilization, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YSW JDD. Performed the experiments: ZWC. Analyzed the data: ZZW. Wrote the paper: ZZW. Took samples: ZWC. Drew the figures: MLW. Revised the manuscript: MLW.