About the Authors

Kira Young

Current address: The Jackson Laboratory, Bar Harbor, Maine 04609, United States of America

Affiliations Center for Molecular Medicine, Maine Medical Center Research Institute, Scarborough, Maine 04074, United States of America, Graduate School of Biomedical Sciences and Engineering, University of Maine, Orono, Maine 04469, United States of America

Eric Tweedie

Affiliation Center for Molecular Medicine, Maine Medical Center Research Institute, Scarborough, Maine 04074, United States of America

Barbara Conley

Affiliation Center for Molecular Medicine, Maine Medical Center Research Institute, Scarborough, Maine 04074, United States of America

Jacquelyn Ames

Affiliations Center for Molecular Medicine, Maine Medical Center Research Institute, Scarborough, Maine 04074, United States of America, Graduate School of Biomedical Sciences and Engineering, University of Maine, Orono, Maine 04469, United States of America

MaryLynn FitzSimons

Affiliation Graduate School of Biomedical Sciences and Engineering, University of Maine, Orono, Maine 04469, United States of America

Peter Brooks

Affiliations Center for Molecular Medicine, Maine Medical Center Research Institute, Scarborough, Maine 04074, United States of America, Graduate School of Biomedical Sciences and Engineering, University of Maine, Orono, Maine 04469, United States of America

Lucy Liaw

Affiliations Center for Molecular Medicine, Maine Medical Center Research Institute, Scarborough, Maine 04074, United States of America, Graduate School of Biomedical Sciences and Engineering, University of Maine, Orono, Maine 04469, United States of America

Calvin P. H. Vary

varyc@mmc.org

Affiliations Center for Molecular Medicine, Maine Medical Center Research Institute, Scarborough, Maine 04074, United States of America, Graduate School of Biomedical Sciences and Engineering, University of Maine, Orono, Maine 04469, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KY BC CPHV. Performed the experiments: KY ET BC MF JA CPHV. Analyzed the data: KY ET BC MF PB LL JA CPHV. Contributed reagents/materials/analysis tools: KY ET BC PB LL JA CPHV. Wrote the paper: KY LL JA CPHV.