About the Authors

Lida Xing

xinglida@gmail.com

Affiliation School of the Earth Sciences and Resources, China University of Geosciences, Beijing 100083, China

Jianping Zhang

Affiliation School of the Earth Sciences and Resources, China University of Geosciences, Beijing 100083, China

Martin G. Lockley

Affiliation Dinosaur Trackers Research Group, University of Colorado Denver, PO Box 173364, Denver, CO 80217, United States of America

Richard T. McCrea

Affiliation Peace Region Palaeontology Research Centre, Box 1540, Tumbler Ridge, British Columbia V0C 2W0, Canada

Hendrik Klein

Affiliation Saurierwelt Paläontologisches Museum, Alte Richt 7, D-92318 Neumarkt, Germany

Luis Alcalá

Affiliation Fundación Conjunto Paleontológico de Teruel–Dinópolis, Teruel, Aragón E-44002, Spain

Lisa G. Buckley

Affiliation Peace Region Palaeontology Research Centre, Box 1540, Tumbler Ridge, British Columbia V0C 2W0, Canada

Michael E. Burns

Affiliation Department of Biological Sciences, University of Alberta, Edmonton, Alberta T6G 2E9, Canada

Susanna B. Kümmell

Affiliation Institute of Evolutionary Biology, University Witten/Herdecke, Stockumerstr. 10–12, 58454 Witten, Germany

Qing He

Affiliation School of the Earth Sciences and Resources, China University of Geosciences, Beijing 100083, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: LX ML JZ. Performed the experiments: LX ML RM HK LB SK QH. Analyzed the data: LX ML RM LB HK QH. Contributed reagents/materials/analysis tools: LX ML RM. Wrote the paper: LX JZ ML RM HK LA LB MB SK QH.