About the Authors

Christopher T. Esapa

Contributed equally to this work with: Christopher T. Esapa, Fadil M. Hannan

Affiliations Academic Endocrine Unit, Radcliffe Department of Medicine, Oxford Centre for Diabetes, Endocrinology and Metabolism (OCDEM), University of Oxford, Oxford, United Kingdom, Medical Research Council (MRC) Mammalian Genetics Unit and Mary Lyon Centre, MRC Harwell, Harwell Science and Innovation Campus, Harwell, United Kingdom

Fadil M. Hannan

Contributed equally to this work with: Christopher T. Esapa, Fadil M. Hannan

Affiliations Academic Endocrine Unit, Radcliffe Department of Medicine, Oxford Centre for Diabetes, Endocrinology and Metabolism (OCDEM), University of Oxford, Oxford, United Kingdom, Department of Musculoskeletal Biology, Institute of Ageing and Chronic Disease, University of Liverpool, Liverpool, United Kingdom

Valerie N. Babinsky

Affiliation Academic Endocrine Unit, Radcliffe Department of Medicine, Oxford Centre for Diabetes, Endocrinology and Metabolism (OCDEM), University of Oxford, Oxford, United Kingdom

Paul Potter

Affiliation Medical Research Council (MRC) Mammalian Genetics Unit and Mary Lyon Centre, MRC Harwell, Harwell Science and Innovation Campus, Harwell, United Kingdom

Gethin P. Thomas

Affiliation University of Queensland Diamantina Institute, Translational Research Institute, Princess Alexandra Hospital, University of Queensland, Brisbane, Australia

Peter I. Croucher

Affiliation Garvan Institute for Medical Research, Sydney, Australia

Matthew A. Brown

Affiliation University of Queensland Diamantina Institute, Translational Research Institute, Princess Alexandra Hospital, University of Queensland, Brisbane, Australia

Steve D. M. Brown

Affiliation Medical Research Council (MRC) Mammalian Genetics Unit and Mary Lyon Centre, MRC Harwell, Harwell Science and Innovation Campus, Harwell, United Kingdom

Roger D. Cox

Affiliation Medical Research Council (MRC) Mammalian Genetics Unit and Mary Lyon Centre, MRC Harwell, Harwell Science and Innovation Campus, Harwell, United Kingdom

Rajesh V. Thakker

rajesh.thakker@ndm.ox.ac.uk

Affiliation Academic Endocrine Unit, Radcliffe Department of Medicine, Oxford Centre for Diabetes, Endocrinology and Metabolism (OCDEM), University of Oxford, Oxford, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CTE FMH VNB PP GPT PIC MAB SDB RDC RVT. Performed the experiments: CTE VNB. Analyzed the data: CTE FMH VNB SDB RDC RVT. Wrote the paper: CTE FMH SDB RDC RVT.