About the Authors

Mari Kekkonen

kekkonen@uoguelph.ca

Affiliations Finnish Museum of Natural History, University of Helsinki, Zoology Unit, University of Helsinki, Helsinki, Finland, Biodiversity Institute of Ontario, University of Guelph, Guelph, Ontario, Canada

Marko Mutanen

Affiliation Department of Genetics and Physiology, University of Oulu, Oulu, Finland

Lauri Kaila

Affiliation Finnish Museum of Natural History, University of Helsinki, Zoology Unit, University of Helsinki, Helsinki, Finland

Marko Nieminen

Affiliation Metapopulation Research Centre, Department of Biosciences, University of Helsinki, Helsinki, Finland

Paul D. N. Hebert

Affiliation Biodiversity Institute of Ontario, University of Guelph, Guelph, Ontario, Canada

Competing Interests

HOK-Elanto has been a sponsor for Finnish Museum of Natural History, sponsoring without specifications activities of the institute. It was an internal decision in the Finnish Museum of Natural History to support the publications in the form of financing a collecting trip of Dr. Lauri Kaila to Australia, with no communication with the sponsor regarding this decision. Therefore, the authors declare impartiality and no influence by HOK-Elanto regarding the support for the activity relating to this publication. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: MK. Performed the experiments: MK MN. Analyzed the data: MK MN. Contributed reagents/materials/analysis tools: PDNH MM LK. Wrote the paper: MK MM LK MN PDNH.