About the Authors

Esther Gramage

Contributed equally to this work with: Esther Gramage, Travis D’Cruz

Affiliation Department of Ophthalmology and Visual Sciences, University of Michigan, Ann Arbor, Michigan, United States of America

Travis D’Cruz

Contributed equally to this work with: Esther Gramage, Travis D’Cruz

Affiliation Department of Ophthalmology and Visual Sciences, University of Michigan, Ann Arbor, Michigan, United States of America

Scott Taylor

Affiliation Department of Ophthalmology and Visual Sciences, University of Michigan, Ann Arbor, Michigan, United States of America

Ryan Thummel

Affiliation Departments of Anatomy and Cell Biology and Ophthalmology, Wayne State University School of Medicine, Detroit, Michigan, United States of America

Peter F. Hitchcock

peterh@med.umich.edu

Affiliation Department of Ophthalmology and Visual Sciences, University of Michigan, Ann Arbor, Michigan, United States of America

Competing Interests

PFH and RT are PLOS ONE Editorial Board members. This does not alter the authors' adherence to PLOS ONE Editorial policies and criteria.

Author Contributions

Conceived and designed the experiments: PFH EG TSD RT. Performed the experiments: EG TSD RT ST. Analyzed the data: PFH EG TSD. Contributed reagents/materials/analysis tools: PFH EG TSD RT ST. Wrote the paper: PFH EG TSD RT ST.