About the Authors

Hyeongmin Kim

‡ Hyeongmin Kim and KDS are joint first authors on this work.

Affiliation Department of Agricultural Biotechnology, Animal Biotechnology Major, and Research Institute for Agriculture and Life Sciences, Seoul National University, Seoul, 151–921, Republic of Korea

Ki Duk Song

‡ Hyeongmin Kim and KDS are joint first authors on this work.

Affiliation Genomic Informatics Center, Hankyong National University, Anseong, 456–749, Republic of Korea

Hyeon Jeong Kim

Affiliation CHO & KIM Genomics, Seoul National University Research Park, Seoul, 151–919, Republic of Korea

WonCheoul Park

Affiliation Department of Agricultural Biotechnology, Animal Biotechnology Major, and Research Institute for Agriculture and Life Sciences, Seoul National University, Seoul, 151–921, Republic of Korea

Jaemin Kim

Affiliation Interdisciplinary Program in Bioinformatics, Seoul National University, Seoul, 151–742, Republic of Korea

Taeheon Lee

Affiliation Department of Agricultural Biotechnology, Animal Biotechnology Major, and Research Institute for Agriculture and Life Sciences, Seoul National University, Seoul, 151–921, Republic of Korea

Dong-Hyun Shin

Affiliation Department of Agricultural Biotechnology, Animal Biotechnology Major, and Research Institute for Agriculture and Life Sciences, Seoul National University, Seoul, 151–921, Republic of Korea

Woori Kwak

Affiliation Interdisciplinary Program in Bioinformatics, Seoul National University, Seoul, 151–742, Republic of Korea

Young-jun Kwon

Affiliation Interdisciplinary Program in Bioinformatics, Seoul National University, Seoul, 151–742, Republic of Korea

Samsun Sung

Affiliation CHO & KIM Genomics, Seoul National University Research Park, Seoul, 151–919, Republic of Korea

Sunjin Moon

Affiliation Department of Agricultural Biotechnology, Animal Biotechnology Major, and Research Institute for Agriculture and Life Sciences, Seoul National University, Seoul, 151–921, Republic of Korea

Kyung-Tai Lee

Affiliation National Institute of Animal Science, RDA, Suwon, 441–706, Republic of Korea

Namshin Kim

Affiliation Korean Bioinformation Center, Korea Research Institute of Bioscience and Biotechnology, Daejeon, 305–806, Republic of Korea

Joon Ki Hong

Affiliation Swine Science Division, National Institute of Animal Science, RDA, Cheonan, 331–801, Republic of Korea

Kyung Yeon Eo

Affiliation Animal Research Division, Seoul Zoo, Seoul, 427–702, Republic of Korea

Kang Seok Seo

Affiliation Department of Animal Science and Technology, College of Life Science and Natural Resources, Sunchon National University, Suncheon, 540–950, Republic of Korea

Girak Kim

Affiliation Department of Agricultural Biotechnology, Animal Biotechnology Major, and Research Institute for Agriculture and Life Sciences, Seoul National University, Seoul, 151–921, Republic of Korea

Sungmoo Park

Affiliation Department of Agricultural Biotechnology, Animal Biotechnology Major, and Research Institute for Agriculture and Life Sciences, Seoul National University, Seoul, 151–921, Republic of Korea

Cheol-Heui Yun

Affiliation Department of Agricultural Biotechnology, Animal Biotechnology Major, and Research Institute for Agriculture and Life Sciences, Seoul National University, Seoul, 151–921, Republic of Korea

Hyunil Kim

Affiliation Optipharm, Inc., 63, Osongsangmyeong 6-ro, Osong-eup, Chengwon-gun, Chungcheongbuk-do, 363–954, Republic of Korea

Kimyung Choi

Affiliation Optipharm, Inc., 63, Osongsangmyeong 6-ro, Osong-eup, Chengwon-gun, Chungcheongbuk-do, 363–954, Republic of Korea

Jiho Kim

Affiliation Optipharm, Inc., 63, Osongsangmyeong 6-ro, Osong-eup, Chengwon-gun, Chungcheongbuk-do, 363–954, Republic of Korea

Woon Kyu Lee

Affiliation Laboratory of Developmental Genetics, College of Medicine, Inha University, Incheon, 400–103, Republic of Korea

Duk-Kyung Kim

Affiliation Genomic Informatics Center, Hankyong National University, Anseong, 456–749, Republic of Korea

Jae-Don Oh

Affiliation Genomic Informatics Center, Hankyong National University, Anseong, 456–749, Republic of Korea

Eui-Soo Kim

Affiliation Department of Animal Sciencs, Iowa State University, Ames, Iowa, 50011, United States of America

Seoae Cho

Affiliation CHO & KIM Genomics, Seoul National University Research Park, Seoul, 151–919, Republic of Korea

Hak-Kyo Lee

breedlee@empas.com (HKL); thkim63@korea.kr (THK); heebal@snu.ac.kr (Heebal Kim)

Affiliation Genomic Informatics Center, Hankyong National University, Anseong, 456–749, Republic of Korea

Tae-Hun Kim

breedlee@empas.com (HKL); thkim63@korea.kr (THK); heebal@snu.ac.kr (Heebal Kim)

Affiliation National Institute of Animal Science, RDA, Suwon, 441–706, Republic of Korea

Heebal Kim

breedlee@empas.com (HKL); thkim63@korea.kr (THK); heebal@snu.ac.kr (Heebal Kim)

Affiliations Department of Agricultural Biotechnology, Animal Biotechnology Major, and Research Institute for Agriculture and Life Sciences, Seoul National University, Seoul, 151–921, Republic of Korea, CHO & KIM Genomics, Seoul National University Research Park, Seoul, 151–919, Republic of Korea

Competing Interests

Authors Hyunil Kim, KC and Jiho Kim are employed by Optipharm, Inc. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: Hyeongmin Kim KDS Heebal Kim. Performed the experiments: WK Hyunil Kim KC Jiho Kim. Analyzed the data: Hyeongmin Kim WCP WK YK SS. Contributed reagents/materials/analysis tools: Jaemin Kim TL DHS SM KTL NK JKH KYE KSS GK SP CHY WKL DKK JDO ESK SC HKL THK. Wrote the paper: Hyeongmin Kim HJK Jaemin Kim.