About the Authors

Norio Takata

takata.norio@keio.jp (NT); kftanaka@keio.jp (KFT)

Affiliations Department of Neuropsychiatry, School of Medicine, Keio University, Shinjuku, Tokyo, Japan, Central Institute for Experimental Animals, Kawasaki, Kanagawa, Japan

Keitaro Yoshida

Contributed equally to this work with: Keitaro Yoshida, Yuji Komaki

Affiliation Department of Neuropsychiatry, School of Medicine, Keio University, Shinjuku, Tokyo, Japan

Yuji Komaki

Contributed equally to this work with: Keitaro Yoshida, Yuji Komaki

Affiliations Department of Physiology, School of Medicine, Keio University, Shinjuku, Tokyo, Japan, Central Institute for Experimental Animals, Kawasaki, Kanagawa, Japan

Ming Xu

Affiliation Department of Neuropsychiatry, School of Medicine, Keio University, Shinjuku, Tokyo, Japan

Yuki Sakai

Affiliations Department of Neuropsychiatry, School of Medicine, Keio University, Shinjuku, Tokyo, Japan, Central Institute for Experimental Animals, Kawasaki, Kanagawa, Japan, Department of Psychiatry, Graduate School of Medicine, Kyoto Prefectural University of Medicine, Kyoto, Japan

Keigo Hikishima

Affiliations Department of Physiology, School of Medicine, Keio University, Shinjuku, Tokyo, Japan, Central Institute for Experimental Animals, Kawasaki, Kanagawa, Japan

Masaru Mimura

Affiliation Department of Neuropsychiatry, School of Medicine, Keio University, Shinjuku, Tokyo, Japan

Hideyuki Okano

Affiliation Department of Physiology, School of Medicine, Keio University, Shinjuku, Tokyo, Japan

Kenji F. Tanaka

takata.norio@keio.jp (NT); kftanaka@keio.jp (KFT)

Affiliation Department of Neuropsychiatry, School of Medicine, Keio University, Shinjuku, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: NT KFT. Performed the experiments: NT KY YK MX. Analyzed the data: NT YK YS. Contributed reagents/materials/analysis tools: KFT KH MM HO. Wrote the paper: NT KFT.