About the Authors

Arata Hidano

Affiliations Viral Disease and Epidemiology Research Division, National Institute of Animal Health, National Agriculture, and Food Research Organization, Tsukuba, Ibaraki, Japan, EpiCentre, Institute of Veterinary, Animal and Biomedical Sciences, Massey University, Palmerston North, New Zealand

Takehisa Yamamoto

Affiliation Viral Disease and Epidemiology Research Division, National Institute of Animal Health, National Agriculture, and Food Research Organization, Tsukuba, Ibaraki, Japan

Yoko Hayama

Affiliation Viral Disease and Epidemiology Research Division, National Institute of Animal Health, National Agriculture, and Food Research Organization, Tsukuba, Ibaraki, Japan

Norihiko Muroga

Affiliation Viral Disease and Epidemiology Research Division, National Institute of Animal Health, National Agriculture, and Food Research Organization, Tsukuba, Ibaraki, Japan

Sota Kobayashi

Affiliation Viral Disease and Epidemiology Research Division, National Institute of Animal Health, National Agriculture, and Food Research Organization, Tsukuba, Ibaraki, Japan

Takeshi Nishida

Affiliation Animal Products Safety Division, Food Safety and Consumer Affairs Bureau, Ministry of Agriculture, Forestry and Fisheries, Chiyoda-ku, Tokyo, Japan

Toshiyuki Tsutsui

tsutsui@affrc.go.jp

Affiliation Viral Disease and Epidemiology Research Division, National Institute of Animal Health, National Agriculture, and Food Research Organization, Tsukuba, Ibaraki, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AH TT. Performed the experiments: AH TT. Analyzed the data: AH. Contributed reagents/materials/analysis tools: AH TY YH NM SK TN TT. Wrote the paper: AH TY YH TT.