About the Authors

Seiya Watanabe

irab@agr.ehime-u.ac.jp

Affiliation Faculty of Agriculture, Ehime University, Matsuyama, Ehime, 790–8566, Japan

Yoshiaki Tanimoto

Affiliation Faculty of Agriculture, Ehime University, Matsuyama, Ehime, 790–8566, Japan

Hisashi Nishiwaki

Affiliation Faculty of Agriculture, Ehime University, Matsuyama, Ehime, 790–8566, Japan

Yasuo Watanabe

Affiliation Faculty of Agriculture, Ehime University, Matsuyama, Ehime, 790–8566, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SW. Performed the experiments: SW YT. Analyzed the data: SW HN. Contributed reagents/materials/analysis tools: SW YW. Wrote the paper: SW.