About the Authors

Satoshi Kume

Affiliation Cellular Function Imaging Team, Division of Bio-function Dynamics Imaging, RIKEN Center for Life Science Technologies, Kobe, Hyogo, Japan

Masanori Yamato

Affiliation Cellular Function Imaging Team, Division of Bio-function Dynamics Imaging, RIKEN Center for Life Science Technologies, Kobe, Hyogo, Japan

Yasuhisa Tamura

Affiliation Cellular Function Imaging Team, Division of Bio-function Dynamics Imaging, RIKEN Center for Life Science Technologies, Kobe, Hyogo, Japan

Guanghua Jin

Affiliation Cellular Function Imaging Team, Division of Bio-function Dynamics Imaging, RIKEN Center for Life Science Technologies, Kobe, Hyogo, Japan

Masayuki Nakano

Affiliations Cellular Function Imaging Team, Division of Bio-function Dynamics Imaging, RIKEN Center for Life Science Technologies, Kobe, Hyogo, Japan, Department of Physiology, Osaka City University Graduate School of Medicine, Osaka, Japan

Yukiharu Miyashige

Affiliation Cellular Function Imaging Team, Division of Bio-function Dynamics Imaging, RIKEN Center for Life Science Technologies, Kobe, Hyogo, Japan

Asami Eguchi

Affiliation Cellular Function Imaging Team, Division of Bio-function Dynamics Imaging, RIKEN Center for Life Science Technologies, Kobe, Hyogo, Japan

Yoshiyuki Ogata

Affiliation Graduate School of Life and Environmental Sciences, Osaka Prefecture University, Osaka, Japan

Nobuhito Goda

Affiliation Department of Life Science and Medical Bio-Science, School of Advanced Science and Engineering, Waseda University, Tokyo, Japan

Kazuhiro Iwai

Affiliation Department of Molecular and Cellular Physiology, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Emi Yamano

Affiliation Department of Physiology, Osaka City University Graduate School of Medicine, Osaka, Japan

Yasuyoshi Watanabe

Affiliations Department of Physiology, Osaka City University Graduate School of Medicine, Osaka, Japan, Pathophysiological and Health Science Team, RIKEN Center for Life Science Technologies, Kobe, Hyogo, Japan

Tomoyoshi Soga

Affiliation Institute for Advanced Biosciences, Keio University, Tsuruoka, Yamagata, Japan

Yosky Kataoka

kataokay@riken.jp

Affiliations Cellular Function Imaging Team, Division of Bio-function Dynamics Imaging, RIKEN Center for Life Science Technologies, Kobe, Hyogo, Japan, Department of Physiology, Osaka City University Graduate School of Medicine, Osaka, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SK YK. Performed the experiments: SK MY YT GJ MN YM AE NG TS YK. Analyzed the data: SK YO NG KI EY YW. Contributed reagents/materials/analysis tools: TS YW YK. Wrote the paper: SK MY YO NG KI EY YW TS YK.