About the Authors

Aaron D. Flesch

flesch@email.arizona.edu

Affiliations School of Natural Resources and the Environment, University of Arizona, Tucson, Arizona, United States of America, Division of Biological Sciences, University of Montana, Missoula, Montana, United States of America

Richard L. Hutto

Affiliation Division of Biological Sciences, University of Montana, Missoula, Montana, United States of America

Willem J. D. van Leeuwen

Affiliation School of Natural Resources and the Environment, University of Arizona, Tucson, Arizona, United States of America

Kyle Hartfield

Affiliation School of Natural Resources and the Environment, University of Arizona, Tucson, Arizona, United States of America

Sky Jacobs

Affiliation Watershed Management Group, Tucson, Arizona, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ADF RLH. Analyzed the data: ADF KH. Contributed reagents/materials/analysis tools: ADF WJDvL KH SJ. Wrote the paper: ADF RLH WJDvL KH SJ. Field Work: ADF SJ.