About the Authors

Maomao Zhang

Affiliations Department of Biology, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America, Cell and Molecular Biology Graduate Group, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Priyanka Kothari

Affiliation Department of Biology, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Michael A. Lampson

lampson@sas.upenn.edu

Affiliations Department of Biology, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America, Cell and Molecular Biology Graduate Group, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MZ MAL. Performed the experiments: MZ PK. Analyzed the data: MZ PK. Contributed reagents/materials/analysis tools: MZ. Wrote the paper: MZ MAL.