About the Authors

Emma K. Bales

‡ These authors contributed equally to this work.

Affiliation Biology Department, James Madison University, Harrisonburg, Virginia, United States of America

Oliver J. Hyman

‡ These authors contributed equally to this work.

Affiliation Biology Department, James Madison University, Harrisonburg, Virginia, United States of America

Andrew H. Loudon

Current address: Department of Zoology, University of British Columbia, Vancouver, BC, Canada

Affiliation Biology Department, James Madison University, Harrisonburg, Virginia, United States of America

Reid N. Harris

Affiliation Biology Department, James Madison University, Harrisonburg, Virginia, United States of America

Gregory Lipps

Current address: Ohio Biodiversity Conservation Partnership, Ohio State University, Columbus, Ohio, United States of America

Affiliation Gregory Lipps, LLC, Toledo, Ohio, United States of America

Eric Chapman

Affiliation Western Pennsylvania Conservancy, Pittsburgh, Pennsylvania, United States of America

Kenneth Roblee

Affiliation New York Department of Environmental Conservation, Albany, New York, United States of America

John D. Kleopfer

Affiliation Virginia Department of Game and Inland Fisheries, Richmond, Virginia, United States of America

Kimberly A. Terrell

terrellk@si.edu

Affiliation Smithsonian Conservation Biology Institute, Front Royal, Virginia, United States of America

Competing Interests

Gregory Lipps is the owner and sole employee of Gregory Lipps, LLC. He has received financial support for his work from the Ohio Division of Wildlife. This does not alter the authors’ adherence to PLOS ONE policies on sharing data and materials since all collected data (with the exception of GPS coordinates of capture sites, which are being withheld to prevent poaching, but available from Kimberly Terrell terrellk@si.edu by request) are available to the scientific community after publication.

Author Contributions

Conceived and designed the experiments: KAT AHL. Performed the experiments: AHL GL KR JDK KAT EC. Analyzed the data: EKB OJH AHL. Contributed reagents/materials/analysis tools: AHL RNH. Wrote the paper: EKB OJH AHL RNH KAT.