About the Authors

Bailey L. Wilberts

Affiliation Department of Veterinary Pathology, College of Veterinary Medicine, Iowa State University, 1600 S. 16th St, Ames, Iowa, 50011, United States of America,

Paulo H. Arruda

Affiliation Department of Veterinary Diagnostic and Production Animal Medicine, College of Veterinary Medicine, Iowa State University, 1600 S. 16th St, Ames, Iowa, 50011, United States of America

Joann M. Kinyon

Affiliation Department of Veterinary Diagnostic and Production Animal Medicine, College of Veterinary Medicine, Iowa State University, 1600 S. 16th St, Ames, Iowa, 50011, United States of America

Tim S. Frana

Affiliation Department of Veterinary Diagnostic and Production Animal Medicine, College of Veterinary Medicine, Iowa State University, 1600 S. 16th St, Ames, Iowa, 50011, United States of America

Chong Wang

Affiliation Department of Veterinary Diagnostic and Production Animal Medicine, College of Veterinary Medicine, Iowa State University, 1600 S. 16th St, Ames, Iowa, 50011, United States of America

Drew R. Magstadt

Affiliation Department of Veterinary Diagnostic and Production Animal Medicine, College of Veterinary Medicine, Iowa State University, 1600 S. 16th St, Ames, Iowa, 50011, United States of America

Darin M. Madson

Affiliation Department of Veterinary Diagnostic and Production Animal Medicine, College of Veterinary Medicine, Iowa State University, 1600 S. 16th St, Ames, Iowa, 50011, United States of America

John F. Patience

Affiliation Department of Animal Science, Iowa State University, 201H Kildee, Ames, Iowa, 50011, United States of America

Eric R. Burrough

burrough@iastate.edu

Affiliation Department of Veterinary Diagnostic and Production Animal Medicine, College of Veterinary Medicine, Iowa State University, 1600 S. 16th St, Ames, Iowa, 50011, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: EB BW D. Madson JP JK TF. Performed the experiments: BW PA D. Magstadt D. Madson EB. Analyzed the data: BW CW EB. Contributed reagents/materials/analysis tools: BW JK TF CW JP. Wrote the paper: BW PA JK TF CW D. Magstadt D. Madson JP EB.