About the Authors

Kristine A. Kuhn

Current address: Division of Rheumatology, University of Colorado School of Medicine, Aurora, Colorado, United States of America

Affiliations Department of Pathology and Immunology, Washington University School of Medicine, St. Louis, Missouri, United States of America, Department of Medicine, Washington University School of Medicine, St. Louis, Missouri, United States of America

Nicholas A. Manieri

Affiliation Department of Pathology and Immunology, Washington University School of Medicine, St. Louis, Missouri, United States of America

Ta-Chiang Liu

Affiliation Department of Pathology and Immunology, Washington University School of Medicine, St. Louis, Missouri, United States of America

Thaddeus S. Stappenbeck

stappenb@wustl.edu

Affiliation Department of Pathology and Immunology, Washington University School of Medicine, St. Louis, Missouri, United States of America

Competing Interests

The authors have read the journal's policy and the authors of this manuscript have the following competing interests: This study was funded in part by Pfizer. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: KAK TSS. Performed the experiments: KAK NAM. Analyzed the data: KAK NAM. Contributed reagents/materials/analysis tools: KAK TCL TSS. Wrote the paper: KAK TSS.