About the Authors

Junye Shen

Contributed equally to this work with: Junye Shen, Chuanli Zhou

Affiliations Key Laboratory of Animal Genetics, Breeding and Reproduction, Ministry of Agricultural, National Engineering Laboratory for Animal Breeding, College of Animal Science and Technology, China Agricultural University, Beijing, P.R. China, Department of Animal Genetics, Breeding and Reproduction, College of Animal Science and Technology, China Agricultural University, Beijing, P.R. China

Chuanli Zhou

Contributed equally to this work with: Junye Shen, Chuanli Zhou

Affiliations Key Laboratory of Animal Genetics, Breeding and Reproduction, Ministry of Agricultural, National Engineering Laboratory for Animal Breeding, College of Animal Science and Technology, China Agricultural University, Beijing, P.R. China, Department of Animal Genetics, Breeding and Reproduction, College of Animal Science and Technology, China Agricultural University, Beijing, P.R. China

Shien Zhu

Affiliation Department of Animal Genetics, Breeding and Reproduction, College of Animal Science and Technology, China Agricultural University, Beijing, P.R. China

Wenqing Shi

Affiliation Animal Husbandry and Veterinary Station of Beijing, Beijing, P.R. China

Maishun Hu

Affiliation Department of Animal Genetics, Breeding and Reproduction, College of Animal Science and Technology, China Agricultural University, Beijing, P.R. China

Xiangwei Fu

Affiliation Department of Animal Genetics, Breeding and Reproduction, College of Animal Science and Technology, China Agricultural University, Beijing, P.R. China

Chuduan Wang

Affiliations Key Laboratory of Animal Genetics, Breeding and Reproduction, Ministry of Agricultural, National Engineering Laboratory for Animal Breeding, College of Animal Science and Technology, China Agricultural University, Beijing, P.R. China, Department of Animal Genetics, Breeding and Reproduction, College of Animal Science and Technology, China Agricultural University, Beijing, P.R. China

Yachun Wang

Affiliations Key Laboratory of Animal Genetics, Breeding and Reproduction, Ministry of Agricultural, National Engineering Laboratory for Animal Breeding, College of Animal Science and Technology, China Agricultural University, Beijing, P.R. China, Department of Animal Genetics, Breeding and Reproduction, College of Animal Science and Technology, China Agricultural University, Beijing, P.R. China

Qin Zhang

Affiliations Key Laboratory of Animal Genetics, Breeding and Reproduction, Ministry of Agricultural, National Engineering Laboratory for Animal Breeding, College of Animal Science and Technology, China Agricultural University, Beijing, P.R. China, Department of Animal Genetics, Breeding and Reproduction, College of Animal Science and Technology, China Agricultural University, Beijing, P.R. China

Ying Yu

yuying@cau.edu.cn

Affiliations Key Laboratory of Animal Genetics, Breeding and Reproduction, Ministry of Agricultural, National Engineering Laboratory for Animal Breeding, College of Animal Science and Technology, China Agricultural University, Beijing, P.R. China, Department of Animal Genetics, Breeding and Reproduction, College of Animal Science and Technology, China Agricultural University, Beijing, P.R. China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YY. Performed the experiments: JYS. Analyzed the data: JYS CLZ YY. Contributed reagents/materials/analysis tools: SEZ WQS MSH XWF CDW YCW QZ YY. Wrote the paper: JYS CLZ YY.