About the Authors

Monika Zavodna

z_monika@hotmail.com

Affiliation Department of Anatomy, University of Otago, Dunedin, New Zealand

Andrew Bagshaw

Affiliation Department of Pathology, University of Otago, Christchurch, New Zealand

Rudiger Brauning

Affiliation AgResearch Limited, Invermay Agricultural Centre, Mosgiel, New Zealand

Neil J. Gemmell

Affiliations Department of Anatomy, University of Otago, Dunedin, New Zealand, Allan Wilson Centre for Molecular Ecology and Evolution, University of Otago, Dunedin, New Zealand

Competing Interests

RB is affiliated to AgResearch Limited and contributed to this study as a consultant. NJG was previously a PLOS ONE Editorial Board member until early 2014. These do not alter the authors' adherence to PLOS ONE Editorial policies and criteria.

Author Contributions

Conceived and designed the experiments: MZ NJG. Performed the experiments: MZ. Analyzed the data: MZ AB RB. Contributed reagents/materials/analysis tools: MZ AB RB NJG. Wrote the paper: MZ AB RB NJG.