About the Authors

Calen P. Ryan

calen.p.ryan@gmail.com

Current address: Laboratory for Human Biology Research, Department of Anthropology, Northwestern University, Evanston, Illinois, United States of America

Affiliation Department of Biological Sciences, University of Manitoba, Winnipeg, Manitoba, Canada

W. Gary Anderson

Affiliation Department of Biological Sciences, University of Manitoba, Winnipeg, Manitoba, Canada

Charlene N. Berkvens

Affiliation Assiniboine Park Zoo, Winnipeg, Manitoba, Canada

James F. Hare

Affiliation Department of Biological Sciences, University of Manitoba, Winnipeg, Manitoba, Canada

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CPR JFH WGA. Performed the experiments: CPR JFH CNB WGA. Analyzed the data: CPR. Contributed reagents/materials/analysis tools: JFA WGA. Contributed to the writing of the manuscript: CPR.