About the Authors

Martin Gilbert

mgilbert@wcs.org

Affiliations Wildlife Conservation Society, Bronx, New York, United States of America, Boyd Orr Centre for Population and Ecosystem Health, Institute of Biodiversity, Animal Health and Comparative Medicine, College of Medical, Veterinary and Life Sciences, University of Glasgow, Glasgow, United Kingdom

Dale G. Miquelle

Affiliation Wildlife Conservation Society, Bronx, New York, United States of America

John M. Goodrich

Affiliation Panthera, New York, New York, United States of America

Richard Reeve

Affiliation Boyd Orr Centre for Population and Ecosystem Health, Institute of Biodiversity, Animal Health and Comparative Medicine, College of Medical, Veterinary and Life Sciences, University of Glasgow, Glasgow, United Kingdom

Sarah Cleaveland

Affiliation Boyd Orr Centre for Population and Ecosystem Health, Institute of Biodiversity, Animal Health and Comparative Medicine, College of Medical, Veterinary and Life Sciences, University of Glasgow, Glasgow, United Kingdom

Louise Matthews

Affiliation Boyd Orr Centre for Population and Ecosystem Health, Institute of Biodiversity, Animal Health and Comparative Medicine, College of Medical, Veterinary and Life Sciences, University of Glasgow, Glasgow, United Kingdom

Damien O. Joly

Affiliations Wildlife Conservation Society, Bronx, New York, United States of America, Metabiota, San Francisco, California, United States of America

Competing Interests

Co-author John Goodrich is a PLOS ONE Editorial Board member. DOJ is currently employed by a commercial company (Metabiota). This does not alter the authors’ adherence to PLOS ONE Editorial policies and criteria.

Author Contributions

Contributed to the writing of the manuscript: MG DGM JMG RR LM SC. Conceived the study concept and objectives: MG DOJ DGM JMG LM SC. Contributed to the design and construction of the model: MG DOJ LM RR.