About the Authors

Tomoaki Ichijo

Affiliations Environmental Science and Microbiology, Graduate School of Pharmaceutical Sciences, Osaka University, Suita, Osaka, Japan, Research Institute for Humanity and Nature, Kyoto, Japan

Yoko Izumi

Affiliation Environmental Science and Microbiology, Graduate School of Pharmaceutical Sciences, Osaka University, Suita, Osaka, Japan

Sayuri Nakamoto

Affiliation Environmental Science and Microbiology, Graduate School of Pharmaceutical Sciences, Osaka University, Suita, Osaka, Japan

Nobuyasu Yamaguchi

yamaguti@phs.osaka-u.ac.jp

Affiliation Environmental Science and Microbiology, Graduate School of Pharmaceutical Sciences, Osaka University, Suita, Osaka, Japan

Masao Nasu

Affiliation Environmental Science and Microbiology, Graduate School of Pharmaceutical Sciences, Osaka University, Suita, Osaka, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TI NY MN. Performed the experiments: TI YI SN. Wrote the paper: TI NY MN.