About the Authors

Soohyun Bang

Contributed equally to this work with: Soohyun Bang, Hyojeong Kwon

Affiliation Department of Biotechnology, Yonsei University, Seoul, Republic of Korea

Hyojeong Kwon

Contributed equally to this work with: Soohyun Bang, Hyojeong Kwon

Affiliation Department of Biotechnology, Yonsei University, Seoul, Republic of Korea

Hyun Sook Hwang

Affiliation Center for Neuro-Medicine, Brain Science Institute, Korea Institute of Science and Technology, Seoul, Republic of Korea

Ki Duk Park

Affiliation Center for Neuro-Medicine, Brain Science Institute, Korea Institute of Science and Technology, Seoul, Republic of Korea

Sung Uk Kim

ysbahn@yonsei.ac.kr (YSB); kimsu@kribb.re.kr (SUK)

Affiliation Industrial Bio-materials Research Center, Korea Research Institute of Bioscience and Biotechnology, Daejeon, Republic of Korea

Yong-Sun Bahn

ysbahn@yonsei.ac.kr (YSB); kimsu@kribb.re.kr (SUK)

Affiliation Department of Biotechnology, Yonsei University, Seoul, Republic of Korea

Competing Interests

Yong-Sun Bahn, the corresponding author of this paper, is a PLOS ONE Editorial Board member. This does not alter the authors’ adherence to PLOS ONE Editorial policies and criteria.

Author Contributions

Conceived and designed the experiments: YSB KDP SUK. Performed the experiments: SB HK HSH. Analyzed the data: SB HK HSH YSB KDP SUK. Contributed reagents/materials/analysis tools: SB HK HSH YSB KDP SUK. Contributed to the writing of the manuscript: SB HK HSH YSB KDP SUK.