About the Authors

Koh Nakamura

Contributed equally to this work with: Koh Nakamura, Shih-Wen Chung

Affiliation Biodiversity Research Center, Academia Sinica, Nangang, Taipei, Taiwan

Shih-Wen Chung

Contributed equally to this work with: Koh Nakamura, Shih-Wen Chung

Affiliation Botanical Garden Division, Taiwan Forestry Research Institute, Taipei, Taiwan

Yoshiko Kono

Affiliation Biodiversity Research Center, Academia Sinica, Nangang, Taipei, Taiwan

Meng-Jung Ho

Affiliation Biodiversity Research Center, Academia Sinica, Nangang, Taipei, Taiwan

Tian-Chuan Hsu

Affiliation Botanical Garden Division, Taiwan Forestry Research Institute, Taipei, Taiwan

Ching-I Peng

bopeng@sinica.edu.tw

Affiliation Biodiversity Research Center, Academia Sinica, Nangang, Taipei, Taiwan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KN SWC YK CIP. Performed the experiments: KN SWC YK MJH. Analyzed the data: KN SWC YK. Contributed reagents/materials/analysis tools: SWC CIP KN TCH. Contributed to the writing of the manuscript: KN SWC YK CIP. Collection of the new species: SWC TCH. Morphological study: KN SWC. Molecular study: KN MJH. Karyological study: YK. Draft preparation: KN.