About the Authors

Nattawadee Nantarat

Affiliations Biological Sciences Program, Faculty of Science, Chulalongkorn University, Bangkok, Thailand, Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, Thailand

Christopher M. Wade

Affiliation School of Life Sciences, University of Nottingham, University Park, Nottingham, United Kingdom

Ekgachai Jeratthitikul

Affiliation Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, Thailand

Chirasak Sutcharit

Affiliation Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, Thailand

Somsak Panha

somsak.pan@chula.ac.th

Affiliation Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, Thailand

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: NN CMW CS SP. Performed the experiments: NN CMW CS SP. Analyzed the data: NN CMW EJ CS SP. Contributed reagents/materials/analysis tools: NN CMW EJ CS SP. Wrote the paper: NN CMW EJ CS SP. Specimens collecting, identification, molecular works (DNA extract, PCR) and manuscript preparation: NN. Morphological and molecular analysis: NN. Taxonomy, morphological and ecological discussion: SP.