About the Authors

Keiji Uchiyama

Contributed equally to this work with: Keiji Uchiyama, Hironori Miyata

Affiliation Division of Molecular Neurobiology, The Institute for Enzyme Research (KOSOKEN), The University of Tokushima, Kuramato, Tokushima, Japan

Hironori Miyata

Contributed equally to this work with: Keiji Uchiyama, Hironori Miyata

Affiliation Animal Research Center, School of Medicine, University of Occupational and Environmental Health, Yahatanishi, Kitakyushu, Japan

Masashi Yano

Affiliation Division of Molecular Neurobiology, The Institute for Enzyme Research (KOSOKEN), The University of Tokushima, Kuramato, Tokushima, Japan

Yoshitaka Yamaguchi

Current address: Department of Food Science, Faculty of Human Life Science, Mimasaka University, Okayama, Japan

Affiliation Division of Molecular Neurobiology, The Institute for Enzyme Research (KOSOKEN), The University of Tokushima, Kuramato, Tokushima, Japan

Morikazu Imamura

Affiliation Influenza and Prion Disease Research Center, National Institute of Animal Health, Tsukuba, Ibaraki, Japan

Naomi Muramatsu

Affiliation Division of Molecular Neurobiology, The Institute for Enzyme Research (KOSOKEN), The University of Tokushima, Kuramato, Tokushima, Japan

Nandita Rani Das

Affiliation Division of Molecular Neurobiology, The Institute for Enzyme Research (KOSOKEN), The University of Tokushima, Kuramato, Tokushima, Japan

Junji Chida

Affiliation Division of Molecular Neurobiology, The Institute for Enzyme Research (KOSOKEN), The University of Tokushima, Kuramato, Tokushima, Japan

Hideyuki Hara

Affiliation Division of Molecular Neurobiology, The Institute for Enzyme Research (KOSOKEN), The University of Tokushima, Kuramato, Tokushima, Japan

Suehiro Sakaguchi

sakaguchi@tokushima-u.ac.jp

Affiliation Division of Molecular Neurobiology, The Institute for Enzyme Research (KOSOKEN), The University of Tokushima, Kuramato, Tokushima, Japan

Competing Interests

The authors declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HS SS. Performed the experiments: KU HS MY YY MI NM NRD JC HH. Analyzed the data: KU HM SS. Wrote the paper: KU SS.