About the Authors

Iain J. T. Thompson

Affiliation Biomedical Sciences Department, Defence Science and Technology Laboratory, Porton Down, Salisbury, Wiltshire, SP4 0JQ, United Kingdom

Elizabeth R. Mann

Affiliation Antigen Presentation Research Group, Imperial College London, Northwick Park and St. Mark's Campus, Watford Road, Harrow, HA1 3UJ, United Kingdom

Margaret G. Stokes

Affiliation Biomedical Sciences Department, Defence Science and Technology Laboratory, Porton Down, Salisbury, Wiltshire, SP4 0JQ, United Kingdom

Nicholas R. English

Affiliation Antigen Presentation Research Group, Imperial College London, Northwick Park and St. Mark's Campus, Watford Road, Harrow, HA1 3UJ, United Kingdom

Stella C. Knight

Affiliation Antigen Presentation Research Group, Imperial College London, Northwick Park and St. Mark's Campus, Watford Road, Harrow, HA1 3UJ, United Kingdom

Diane Williamson

dewilliamson@dstl.gov.uk

Affiliation Biomedical Sciences Department, Defence Science and Technology Laboratory, Porton Down, Salisbury, Wiltshire, SP4 0JQ, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: IJTT ERM NRE SCK DW. Performed the experiments: IJTT ERM NRE MGS DW. Analyzed the data: IJTT ERM SCK DW. Contributed reagents/materials/analysis tools: NRE ERM IJTT. Wrote the paper: IJTT REM SCK DW.