About the Authors

Qisheng Zuo

‡ QSZ and DL contributed equally to this work.

Affiliation Key Laboratory of Animal Breeding Reproduction and Molecular Design for Jiangsu Province, College of Animal Science and Technology, Yangzhou University, Yangzhou, 225009, China

Dong Li

‡ QSZ and DL contributed equally to this work.

Affiliation Key Laboratory of Animal Breeding Reproduction and Molecular Design for Jiangsu Province, College of Animal Science and Technology, Yangzhou University, Yangzhou, 225009, China

Lei Zhang

Affiliation Key Laboratory of Animal Breeding Reproduction and Molecular Design for Jiangsu Province, College of Animal Science and Technology, Yangzhou University, Yangzhou, 225009, China

Ahmed Kamel Elsayed

Affiliations Key Laboratory of Animal Breeding Reproduction and Molecular Design for Jiangsu Province, College of Animal Science and Technology, Yangzhou University, Yangzhou, 225009, China, College of Veterinary medicine, Suez Canal University, Ismailia, 41522, Egypt

Chao Lian

Affiliation Key Laboratory of Animal Breeding Reproduction and Molecular Design for Jiangsu Province, College of Animal Science and Technology, Yangzhou University, Yangzhou, 225009, China

Qingqing Shi

Affiliation Key Laboratory of Animal Breeding Reproduction and Molecular Design for Jiangsu Province, College of Animal Science and Technology, Yangzhou University, Yangzhou, 225009, China

Zhentao Zhang

Affiliation Key Laboratory of Animal Breeding Reproduction and Molecular Design for Jiangsu Province, College of Animal Science and Technology, Yangzhou University, Yangzhou, 225009, China

Rui Zhu

Affiliation Key Laboratory of Animal Breeding Reproduction and Molecular Design for Jiangsu Province, College of Animal Science and Technology, Yangzhou University, Yangzhou, 225009, China

Yinjie Wang

Affiliation Key Laboratory of Animal Breeding Reproduction and Molecular Design for Jiangsu Province, College of Animal Science and Technology, Yangzhou University, Yangzhou, 225009, China

Kai Jin

Affiliation Key Laboratory of Animal Breeding Reproduction and Molecular Design for Jiangsu Province, College of Animal Science and Technology, Yangzhou University, Yangzhou, 225009, China

Yani Zhang

ynzhang@yzu.edu.cn (YNZ); yubcli@yzu.edu.cn (BCL)

Affiliation Key Laboratory of Animal Breeding Reproduction and Molecular Design for Jiangsu Province, College of Animal Science and Technology, Yangzhou University, Yangzhou, 225009, China

Bichun Li

ynzhang@yzu.edu.cn (YNZ); yubcli@yzu.edu.cn (BCL)

Affiliation Key Laboratory of Animal Breeding Reproduction and Molecular Design for Jiangsu Province, College of Animal Science and Technology, Yangzhou University, Yangzhou, 225009, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BCL. Performed the experiments: QSZ QQS ZTZ. Analyzed the data: QSZ DL. Wrote the paper: QZS YNZ. Edited the paper: QZS LZ RZ KJ YJW CL AKE.