About the Authors

Emiko Yoda

Affiliation Division of Health Chemistry, Department of Healthcare and Regulatory Sciences, School of Pharmacy, Showa University, Shinagawa-ku, Tokyo, Japan

Kohmi Rai

Affiliation Division of Health Chemistry, Department of Healthcare and Regulatory Sciences, School of Pharmacy, Showa University, Shinagawa-ku, Tokyo, Japan

Mai Ogawa

Affiliation Division of Health Chemistry, Department of Healthcare and Regulatory Sciences, School of Pharmacy, Showa University, Shinagawa-ku, Tokyo, Japan

Yuki Takakura

Affiliation Division of Health Chemistry, Department of Healthcare and Regulatory Sciences, School of Pharmacy, Showa University, Shinagawa-ku, Tokyo, Japan

Hiroshi Kuwata

Affiliation Division of Health Chemistry, Department of Healthcare and Regulatory Sciences, School of Pharmacy, Showa University, Shinagawa-ku, Tokyo, Japan

Hidenori Suzuki

Affiliation Division of Morphological and Biomolecular Research, Graduate School of Medicine, Nippon Medical School, Bunkyo-ku, Tokyo, Japan

Yoshihito Nakatani

Affiliation Division of Health Chemistry, Department of Healthcare and Regulatory Sciences, School of Pharmacy, Showa University, Shinagawa-ku, Tokyo, Japan

Makoto Murakami

Affiliation Lipid Metabolism Project, Tokyo Metropolitan Institute of Medical Science, Setagaya-ku, Tokyo, Japan

Shuntaro Hara

haras@pharm.showa-u.ac.jp

Affiliation Division of Health Chemistry, Department of Healthcare and Regulatory Sciences, School of Pharmacy, Showa University, Shinagawa-ku, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: EY HS MM SH. Performed the experiments: EY KR MO YT HS. Analyzed the data: EY HK HS YN MM SH. Contributed reagents/materials/analysis tools: HS HK MM. Contributed to the writing of the manuscript: EY MM SH.