About the Authors

Sanne Terryn

Affiliations National Reference Centre of Rabies, Viral Diseases, Scientific Institute of Public Health (WIV-ISP), Brussels, Belgium, Laboratory of Virology, Department of Virology, Parasitology and Immunology, Faculty of Veterinary Medicine, Ghent University, Merelbeke, Belgium

Aurélie Francart

Affiliation National Reference Centre of Rabies, Viral Diseases, Scientific Institute of Public Health (WIV-ISP), Brussels, Belgium

Sophie Lamoral

Affiliation National Reference Centre of Rabies, Viral Diseases, Scientific Institute of Public Health (WIV-ISP), Brussels, Belgium

Anna Hultberg

Affiliation Ablynx NV, Gent, Belgium

Heidi Rommelaere

Affiliation Ablynx NV, Gent, Belgium

Angela Wittelsberger

Affiliation Ablynx NV, Gent, Belgium

Filip Callewaert

Affiliation Ablynx NV, Gent, Belgium

Thomas Stohr

Affiliation Ablynx NV, Gent, Belgium

Kris Meerschaert

Affiliation Ablynx NV, Gent, Belgium

Ingrid Ottevaere

Affiliation Ablynx NV, Gent, Belgium

Catelijne Stortelers

Affiliation Ablynx NV, Gent, Belgium

Peter Vanlandschoot

Affiliation Ablynx NV, Gent, Belgium

Michael Kalai

Affiliation National Reference Centre of Rabies, Viral Diseases, Scientific Institute of Public Health (WIV-ISP), Brussels, Belgium

Steven Van Gucht

Steven.VanGucht@wiv-isp.be

Affiliations National Reference Centre of Rabies, Viral Diseases, Scientific Institute of Public Health (WIV-ISP), Brussels, Belgium, Laboratory of Virology, Department of Virology, Parasitology and Immunology, Faculty of Veterinary Medicine, Ghent University, Merelbeke, Belgium

Competing Interests

The authors of this manuscript have the following competing interests: HR, FC, TS, KM, IO and CS are employees of Ablynx NV. AH, AW and PV are former employees of Ablynx NV. Ablynx NV produced the VHH and provided the necessary materials, whereas all virus experiments were performed independently at the Scientific Institute of Public Health (WIV-ISP, a government-funded institute). This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: ST AH HR AW CS PV SVG. Performed the experiments: ST AF SL AH HR AW FC TS KM IO CS. Analyzed the data: ST AH HR AW FC TS KM IO CS PV MK SVG. Wrote the paper: ST HR FC TS KM IO CS PV MK SVG.