About the Authors

Shuichi Shibuya

Affiliation Department of Advanced Aging Medicine, Chiba University Graduate School of Medicine, Chiba, Japan

Yusuke Ozawa

Affiliation Department of Advanced Aging Medicine, Chiba University Graduate School of Medicine, Chiba, Japan

Kenji Watanabe

Affiliation Department of Advanced Aging Medicine, Chiba University Graduate School of Medicine, Chiba, Japan

Naotaka Izuo

Affiliation Department of Advanced Aging Medicine, Chiba University Graduate School of Medicine, Chiba, Japan

Toshihiko Toda

Affiliation Department of Advanced Aging Medicine, Chiba University Graduate School of Medicine, Chiba, Japan

Koutaro Yokote

Affiliation Department of Clinical Cell Biology and Medicine, Chiba University Graduate School of Medicine, Chiba, Japan

Takahiko Shimizu

shimizut@chiba-u.jp

Affiliations Department of Advanced Aging Medicine, Chiba University Graduate School of Medicine, Chiba, Japan, Molecular Gerontology, Tokyo Metropolitan Institute of Gerontology, Itabashi, Tokyo, Japan

Competing Interests

Musashino Pharmaceutical Company made a grant donation to Department of Advanced Aging Medicine, Chiba University Graduate School of Medicine, for the conduct of the animal study and for the purchase of reagents. Musashino Pharmaceutical was not involved with the scientific content of the research projects run by the Department of Advanced Aging Medicine, Chiba University Graduate School of Medicine. The authors were not paid employees and did not receive any kind of fees from Musashino Pharmaceutical Company. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: SS TS. Performed the experiments: SS YO KW NI TT. Analyzed the data: SS TS. Wrote the paper: SS KY TS.