About the Authors

Sudarath Baicharoen

Affiliations Bioscience Program, Faculty of Science, Kasetsart University, Bangkok, Thailand, Center for Advanced Studies in Tropical Natural Resources, National Research University-Kasetsart University, Bangkok, Thailand, Conservation, Research and Education Division, Zoological Park Organization, Bangkok, Thailand

Takako Miyabe-Nishiwaki

Affiliation Primate Research Institute, Kyoto University, Inuyama, Aichi, Japan

Visit Arsaithamkul

Affiliation Conservation, Research and Education Division, Zoological Park Organization, Bangkok, Thailand

Yuriko Hirai

Affiliation Primate Research Institute, Kyoto University, Inuyama, Aichi, Japan

Kwanruen Duangsa-ard

Affiliation Chiang Mai Zoo, Chiang Mai, Thailand

Boripat Siriaroonrat

Affiliation Conservation, Research and Education Division, Zoological Park Organization, Bangkok, Thailand

Hiroshi Domae

Affiliation Ishikawa Zoo, Tokusan-machi, Noumi, Ishikawa, Japan

Kornsorn Srikulnath

Affiliations Center for Advanced Studies in Tropical Natural Resources, National Research University-Kasetsart University, Bangkok, Thailand, Department of Genetics, Faculty of Science, Kasetsart University, Bangkok, Thailand

Akihiko Koga

Affiliation Primate Research Institute, Kyoto University, Inuyama, Aichi, Japan

Hirohisa Hirai

hirohisa.hirai.7w@kyoto-u.ac.jp

Affiliation Primate Research Institute, Kyoto University, Inuyama, Aichi, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SB BS VA KS AK HH. Performed the experiments: SB YH AK HH. Analyzed the data: SB YH AK HH. Contributed reagents/materials/analysis tools: SB TMN VA YH KD BS HD KS AK HH. Contributed to the writing of the manuscript: SB TMN AK HH.