About the Authors

James L. Rainford

jlr516@york.ac.uk

Affiliation Department of Biology, University of York, York, United Kingdom

Michael Hofreiter

Affiliations Department of Biology, University of York, York, United Kingdom, Faculty of Mathematics and Natural Sciences, Institute for Biochemistry and Biology, University of Potsdam, Potsdam, Germany

David B. Nicholson

Affiliations Department of Biology, University of York, York, United Kingdom, Department of Natural Sciences, National Museums Scotland, Edinburgh, United Kingdom, Department of Earth Sciences, The Natural History Museum, London, United Kingdom

Peter J. Mayhew

Affiliation Department of Biology, University of York, York, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JR MH PM. Performed the experiments: JR. Analyzed the data: JR. Contributed reagents/materials/analysis tools: JR DN. Wrote the paper: JR MH DN PM.