About the Authors

Ryutaro Kimura

Affiliation Graduate School of Life Science, Hokkaido University, Sapporo, Hokkaido, Japan

Chie Murata

Current address: Department of Human Genetics, Institute of Health Biosciences, the University of Tokushima Graduate School, Tokushima, Tokushima, Japan

Affiliation Graduate School of Life Science, Hokkaido University, Sapporo, Hokkaido, Japan

Yoko Kuroki

Current address: Department of Integrative Genomics, Tohoku Medical Megabank Organization, Tohoku University, Sendai, Miyagi, Japan

Affiliation RIKEN, Center for Integrative Medical Sciences, Yokohama, Kanagawa, Japan

Asato Kuroiwa

asatok@sci.hokudai.ac.jp

Affiliations Graduate School of Life Science, Hokkaido University, Sapporo, Hokkaido, Japan, Laboratory of Animal Cytogenetics, Department of Biological Sciences, Faculty of Science, Hokkaido University, Sapporo, Hokkaido, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AK. Performed the experiments: RK. Analyzed the data: RK AK. Contributed reagents/materials/analysis tools: CM YK. Wrote the paper: AK RK.