About the Authors

Justin J. Greenlee

justin.greenlee@ars.usda.gov

Affiliation Virus and Prion Research Unit, National Animal Disease Center, Agricultural Research Service, United States Department of Agriculture, Ames, Iowa, United States of America

Robert A. Kunkle

Affiliation Virus and Prion Research Unit, National Animal Disease Center, Agricultural Research Service, United States Department of Agriculture, Ames, Iowa, United States of America

Jürgen A. Richt

Current address: Department of Diagnostic Medicine/Pathobiology, College of Veterinary Medicine, Kansas State University, Manhattan, Kansas, United States of America

Affiliation Virus and Prion Research Unit, National Animal Disease Center, Agricultural Research Service, United States Department of Agriculture, Ames, Iowa, United States of America

Eric M. Nicholson

Affiliation Virus and Prion Research Unit, National Animal Disease Center, Agricultural Research Service, United States Department of Agriculture, Ames, Iowa, United States of America

Amir N. Hamir

† Deceased.

Affiliation Virus and Prion Research Unit, National Animal Disease Center, Agricultural Research Service, United States Department of Agriculture, Ames, Iowa, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RAK JAR ANH. Performed the experiments: JJG RAK JAR EMN ANH. Analyzed the data: JJG RAK JAR EMN ANH. Contributed to the writing of the manuscript: JJG RAK. Critical revision of the manuscript: JAR EMN.