About the Authors

Qingran Kong

Contributed equally to this work with: Qingran Kong, Tang Hai

Affiliation Laboratory of Embryo Biotechnology, College of life science, Northeast Agricultural University, Harbin, China

Tang Hai

Contributed equally to this work with: Qingran Kong, Tang Hai

Affiliation State Key Laboratory of Reproductive Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, China

Jing Ma

Affiliation Laboratory of Embryo Biotechnology, College of life science, Northeast Agricultural University, Harbin, China

Tianqing Huang

Affiliation Laboratory of Embryo Biotechnology, College of life science, Northeast Agricultural University, Harbin, China

Dandan Jiang

Affiliation Laboratory of Embryo Biotechnology, College of life science, Northeast Agricultural University, Harbin, China

Bingteng Xie

Affiliation Laboratory of Embryo Biotechnology, College of life science, Northeast Agricultural University, Harbin, China

Meiling Wu

Affiliation Laboratory of Embryo Biotechnology, College of life science, Northeast Agricultural University, Harbin, China

Jiaqiang Wang

Affiliation Laboratory of Embryo Biotechnology, College of life science, Northeast Agricultural University, Harbin, China

Yuran Song

Affiliations State Key Laboratory of Reproductive Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, China, Graduate University of the Chinese Academy of Sciences, Beijing, China

Ying Wang

Affiliation State Key Laboratory of Reproductive Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, China

Yilong He

Affiliation Laboratory of Embryo Biotechnology, College of life science, Northeast Agricultural University, Harbin, China

Jialu Sun

Affiliation Laboratory of Embryo Biotechnology, College of life science, Northeast Agricultural University, Harbin, China

Kui Hu

Affiliation Laboratory of Embryo Biotechnology, College of life science, Northeast Agricultural University, Harbin, China

Runfa Guo

Affiliation Laboratory of Embryo Biotechnology, College of life science, Northeast Agricultural University, Harbin, China

Liu Wang

Affiliation State Key Laboratory of Reproductive Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, China

Qi Zhou

Affiliation State Key Laboratory of Reproductive Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, China

Yanshuang Mu

liu086@126.com (YSM); muyanshuang@163.com (ZHL)

Affiliation Laboratory of Embryo Biotechnology, College of life science, Northeast Agricultural University, Harbin, China

Zhonghua Liu

liu086@126.com (YSM); muyanshuang@163.com (ZHL)

Affiliation Laboratory of Embryo Biotechnology, College of life science, Northeast Agricultural University, Harbin, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: QRK TH YSM JM TQH. Performed the experiments: QRK TH DDJ BTX MLW JQW KH RFG. Analyzed the data: YRS YW YLH JLS. Contributed reagents/materials/analysis tools: ZHL. Contributed to the writing of the manuscript: QRK QZ ZHL.