About the Authors

Liang Meng

Affiliations College of Marine Life Science, Ocean University of China, Qingdao, PR China, Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, PR China

Ying Zhu

Affiliation Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, PR China

Ning Zhang

Affiliation Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, PR China

Wanjun Liu

Affiliation Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, PR China

Yang Liu

Affiliation Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, PR China

Changwei Shao

Affiliation Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, PR China

Na Wang

Affiliation Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, PR China

Songlin Chen

chensl@ysfri.ac.cn

Affiliation Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, PR China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experients: SC LM. Performed the experiments: LM YZ NZ WL YL CS. Analyzed the data: LM YZ NZ. Contributed reagents/materials/analysis tools: SC LM NW. Wrote the manuscript: LM. Revised the manuscript: SC.