About the Authors

Ellen J. Bennett

Affiliation Sheffield Institute for Translational Neuroscience, Department of Neuroscience, University of Sheffield, Sheffield, United Kingdom

Richard J. Mead

Affiliation Sheffield Institute for Translational Neuroscience, Department of Neuroscience, University of Sheffield, Sheffield, United Kingdom

Mimoun Azzouz

Affiliation Sheffield Institute for Translational Neuroscience, Department of Neuroscience, University of Sheffield, Sheffield, United Kingdom

Pamela J. Shaw

Affiliation Sheffield Institute for Translational Neuroscience, Department of Neuroscience, University of Sheffield, Sheffield, United Kingdom

Andrew J. Grierson

a.j.grierson@sheffield.ac.uk

Affiliation Sheffield Institute for Translational Neuroscience, Department of Neuroscience, University of Sheffield, Sheffield, United Kingdom

Competing Interests

Co-author Andrew Grierson is a PLOS ONE Editorial Board member. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: EJB RJM PJS AJG. Performed the experiments: EJM RJM AJG. Analyzed the data: EJB RJM AJG. Wrote the paper: AJG. Revised the manuscript: MA RJM EJB PJS.