About the Authors

Michela Balestri

Affiliations Department of Social Sciences, Oxford Brookes University, Oxford, United Kingdom, Department of Biology, University of Pisa, Pisa, Italy

Marta Barresi

Affiliation Department of Biology, University of Pisa, Pisa, Italy

Marco Campera

Affiliations Department of Social Sciences, Oxford Brookes University, Oxford, United Kingdom, Department of Biology, University of Pisa, Pisa, Italy

Valentina Serra

Affiliation Department of Biology, University of Pisa, Pisa, Italy

Jean Baptiste Ramanamanjato

Affiliation QIT Madagascar Minerals, Rio Tinto, Tolagnaro, Madagascar

Michael Heistermann

Affiliation Endocrinology Laboratory, German Primate Center, Leibniz Institute for Primate Research, Goettingen, Germany

Giuseppe Donati

gdonati@brookes.ac.uk

Affiliation Department of Social Sciences, Oxford Brookes University, Oxford, United Kingdom

Competing Interests

The authors declare that the affiliation of one of them to a commercial company (QIT Madagascar Minerals) does not alter their adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: GD MH. Performed the experiments: M. Barresi M. Balestri MC VS. Analyzed the data: M. Barresi M. Balestri MC VS MH GD. Contributed reagents/materials/analysis tools: JBR MH GD. Wrote the paper: M. Balestri MC JBR MH GD.