About the Authors

Victoria L. Newton

Affiliation Faculty of Life Sciences, University of Manchester, Manchester, United Kingdom

Sumia Ali

Affiliation Faculty of Life Sciences, University of Manchester, Manchester, United Kingdom

Graham Duddy

Current address: The Wellcome Trust Sanger Institute, Hinxton, Cambridge, United Kingdom

Affiliation Platform Technology and Sciences, GlaxoSmithKline, Stevenage, Herts, United Kingdom

Alan J. Whitmarsh

Affiliation Faculty of Life Sciences, University of Manchester, Manchester, United Kingdom

Natalie J. Gardiner

natalie.gardiner@manchester.ac.uk

Affiliation Faculty of Life Sciences, University of Manchester, Manchester, United Kingdom

Competing Interests

This work was supported by a GlaxoSmithKline(GSK)-BBSRC(UK) PhD studentship (VLN). The authors thank GSK for advice, the supply of ASK1n mice and consumable support. This does not alter the authors' adherence to PLOS ONE policies on sharing data. The ASK1 transgenic mice were generated using patented technologies and as a result of licence restrictions are not available to non-collaborators.

Author Contributions

Conceived and designed the experiments: NJG VLN SA AJW. Performed the experiments: VLN SA. Analyzed the data: VLN SA. Contributed reagents/materials/analysis tools: GD. Wrote the paper: VLN SA NJG.